Após a saída de Brenda Leigh Johnson, a capitã Sharon Raydor assume a divisão de crimes da polícia de Los Angeles, mas enfrenta a rejeição da equipe que teve problemas com ela anteriormente.
The squad scrambles to garner a confession when the pregnant wife of a big-time film director is found murdered. Meanwhile, just as things seem to fall into place under Raydor’s leadership, new Deputy D.A. Emma Rios challenges Raydor’s intentions as Rusty’s guardian.
A complex murder-suicide case has the department reeling. Meanwhile, when Rusty receives a mysterious and threatening letter Emma demands that he be relocated. Torn between her desire to protect Rusty and professional duty to the case, Raydor is forced to seriously consider Emma’s request.
A meddling television writer-producer accompanies Sanchez and Tao on a ride-along so he can get an inside look at the LAPD. But what starts out as a simple outing quickly gets out of hand. And Rusty employs some unfair help when his teacher makes him re-write an essay.
The LAPD flies a material witness into Los Angeles to testify, but in a horrible turn of events their witness winds up as the key suspect in an entirely new murder. Personal and professional stress takes a toll on Flynn. And the squad takes bets when Rusty brings a girl from class to hang out with him at the department.
The squad investigates a brutal murder, but they soon find that dark secrets are at the root of an apparent drug bust gone wrong. Meanwhile, Raydor’s estranged husband of 25 years returns to Los Angeles to work as a court-appointed attorney. Charming and charismatic, Jack becomes fast friends with Rusty but Sharon continues to remain wary.
When a child with gender dysphoria goes missing, the Major Crimes division scrambles to solve the emotional case before it's too late. Torn between several suspects, the squad has to be careful as it tries to connect the evidence to a dangerous bully. Meanwhile, Jack Raydor attempts to make up for years of lousy parenting.
In a case that brings gang violence and social profiling to the forefront, the squad realizes that things are not always as they seem. Complicating matters further, Sharon offers Jack the opportunity to be a court-appointed attorney in the case. Meanwhile, Rusty expresses apprehension about meeting Kris' parents.
The media goes into a frenzy over race relations when the body of a young Latino man is found at the home of a white award-winning swimming coach. Flynn struggles with an emotional dilemma on the eve of his daughter's wedding. Kris' mother pays Raydor a visit to discuss some concerns.
Provenza faces possible permanent desk duty while the squad investigates the suspicious death of a loathsome ICE agent which brings them into contact with the residents of an ancient apartment complex nicknamed La Shangri-la.
In a case that hits a little too close to home for Rusty, a 19-year-old prostitute is brutally murdered. The squad rushes to reach a deal with the remorseless killer they believe is part of a larger criminal enterprise. Their efforts, however, do not sit well with the judge assigned to the case.
The team works strategically to capture a spree killer who has put everyone on high alert. Trying to solve the case quickly, Raydor is torn between two seemingly right decisions, each of which comes with major consequences. Meanwhile, Rusty is given some extra freedom, but more threats leave him afraid to stray far from home.
After a bad batch of drugs claims the lives of two brothers, the race is on for the squad to find the dealer before more of their fellow high school students are threatened. Feeling smothered by his new around-the-clock security detail, Rusty faces a big decision.
When a newly released prisoner misses two back-to-back parole meetings, the squad is called in to track him down after the suspect’s therapist expresses concerns over his mental state. Meanwhile, Sharon invites therapist Dr. Joe Bowman to the office to evaluate Rusty over a game of chess.
When a body turns up at a posh country club in a quiet neighborhood, the squad’s investigation leads them down a money trail full of surprises. Flynn is noticeably distracted on the job, and Emma’s actions have Rusty questioning her intentions, causing him to distrust her even more.
Christmas vacation is interrupted for the Major Crimes Division when a baseball scout is found dead in his RV. The squad works through the murder case, unfolding a complicated kidnapping scheme. Meanwhile, Rusty opens up to Dr. Joe about his mom.
When a well-off graduate student gets shot to death in a gang-riddled neighborhood, the Major Crimes Division must work with the FBI, including Fritz Howard, to find his killer. Throughout the investigation, it is revealed how each member of the unit became interested in joining the police force.
A year-end blowout sale becomes literal for a used-car lot after one of its owners is killed when a bomb explodes. While the squad tries to find who is responsible, Rusty dons a bullet-proof vest under the watchful eye of Lt. Chuck Cooper, an SIS (Special Investigation Section) supervisor, in an attempt to catch the person responsible for threatening his and Sharon’s life.
The Major Crimes unit draws closer to finding a dangerous serial killer. Rusty faces his biggest challenge yet when he bravely takes the stand against Phillip Stroh. Complicating the case further, the team goes head to head with Linda Rothman, a tough, powerful attorney who doesn't let morals stand in her way of winning a case.
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